1. University of Bristol, School of Veterinary Medicine, Clinical Veterinary Science, Langford, Bristol, BS 18 7DU, UK

2. Centre for Tropical Veterinary Medicine, University of Edinburgh, Easter Bush, Roslin, Midlothian, EH25 9RG, UK

3. University of Glasgow, Anderson College, Department of Molecular Genetics, 56 Dumbarton Road, Glasgow, G1 6NU UK

4. University of Liverpool, Faculty of Veterinary Science, Department of Clinical Veterinary Science and Animal Husbandry, Leahurst, Neston, South Wirral, L64 7TE, U.K